Beginning in the early 1990s, a few hominid remains were found, but the most informative specimens are three well-preserved crania, the most recently discovered (in 2001) being almost complete (see photos). These remains (all dated to approximately 1.8 m.y.a.) are important because they are the best preserved hominids of this antiquity found anywhere outside of Africa. Moreover, they show a mixed pattern of characteristics, some quite unexpected.
In some respects the Dmanisi crania are similar to H. erectus (for example, the long, low vault, wide base, and thickening along the sagittal midline). In other characteristics, however, the Dmanisi individuals are different from other hominid finds outside of Africa. In particular, the most complete specimen (No. 2700) has a less robust and thinner browridge, a projecting lower face, and a large upper canine. Thus, at least from the front, this skull is highly reminiscent of the smaller early Homo specimens from East Africa more so than of Homo erectus. Moreover, cranial capacity if this individual is very small (estimated at only 600 cm3 , well within the range of early Homo). In fact, all three Dmanisi crania show small cranial capacities (the other two estimated at 650 and 780 cm3).
|Homo erectus from Dmanisi, Republic of Georgia
Photos courtesy of Professor David Lordkipanidze, Deputy Director, Georgian State Museum
A number of stone tools have also been recovered at Dmanisi. The tools are similar to early African implements and are quite different from the ostensibly more advanced technology of the Acheulian (the latter broadly associated with H. erectus in much of the Old World).
From these recent, somewhat startling revelations from Dmanisi several questions can be raised:
1) Was Homo erectus the first hominid to leave Africa -- or was it an earlier form of Homo?
2) Did hominids require a large brain and sophisticated stone-tool culture to disperse out of Africa?
3) Was the large, robust body build of H. erectus a necessary adaptation to disperse initially into Eurasia.
4) Did, in fact, H. erectus evolve primarily in Eurasia and then migrate back to Africa?
Of course, the Dmanisi discoveries are very new, so any conclusions we draw must be seen as highly tentative. Nevertheless, the recent evidence raises important and exciting possibilities. In regards to Question 1, it now seems likely the first trans-continental hominid migrants were a form of early Homo (similar to the smaller East African species, Homo habilis). At best, and as exemplified at Dmanisi, the first hominids to leave Africa were a very early form of H. erectus, one much more primitive than any of the other specimens from Africa, Asia, or Europe discussed above.
As for question 2, certainly the smaller individuals from Dmanisi did not have a large brain (by H. erectus standards), nor did they have an advanced stone tool culture (possessing tools very similar to the earliest ones from East and South Africa).
Question 3 concerns body size and proportions, but is more speculative than the two above queries. Very little postcranial material has been found thus far at Dmanisi, so we do not know as yet the body structure of the earliest hominids to leave Africa. It is possible, however, that the overall body proportions (as with the face of the smallest Dmanisi cranium) resemble Homo habilis more than they do H. erectus. Thus, these first pioneers to leave Africa may have been, in the words of Phillip Rightmire (of the University of Binghamton and a co-author of the first article announcing the 2001 Dmanisi discovery), ‘little people.” That is, they may have been very different from the big-bodied, long-legged, full-blown H. erectus body plan.
And, lastly, where did the full-blown H. erectus morphology first evolve? For decades it has been widely assumed H. erectus evolved first in Africa and then emigrated to elsewhere in the Old World. Hence, now the teasing possibility exists that H. erectus evolved primarily in Eurasia and (only after attaining its fully characteristic morphology) did it migrate back into Africa (as perhaps the Bouri cranium, dated to 1 m.y.a. suggests?).
by Robert Jurmain, San Jose State University
Georgia became a kingdom about 4 B.C. and Christianity was introduced in A.D. 337. During the reign of Queen Tamara (1184–1213), its territory included the whole of Transcaucasia. During the 13th century, Tamerlane and the Mongols decimated its population. From the 16th century on, the country was the scene of a struggle between Persia and Turkey. In the 18th century, it became a vassal to Russia in exchange for protection from the Turks and Persians.
Georgia joined Azerbaijan and Armenia in 1917 to establish the anti-Bolshevik Transcaucasian Federation and on its dissolution in 1918 proclaimed its independence. In 1922, Georgia, Armenia, and Azerbaijan were annexed by the USSR and formed the Transcaucasian Soviet Socialist Republic. In 1936, Georgia became a separate Soviet republic. Under Soviet rule it was transformed from an agrarian country to a largely industrial urban society.
Georgia proclaimed its independence from the USSR on April 6, 1991. In Jan. 1992, its leader, Zviad Gamsakhurdia, was sacked and later accused of dictatorial policies, the jailing of opposition leaders, human rights abuses, and clamping down on the media. A ruling military council was established by the opposition until a civilian authority could be restored. In 1992, Eduard Shevardnadze, the Soviet Union's foreign minister under Gorbachev, became president.
In 1992–1993, the government engaged in armed conflict with separatists in the breakaway province of Abkhazia. In 1994, Russia and Georgia signed a cooperation treaty that authorized Russia to keep three military bases in Georgia and allowed Russians to train and equip the Georgian army. In 1996, Georgia and its breakaway region of South Ossetia agreed to a cessation of hostilities in their six-year conflict. With little progress in resolving the Abkhazia situation, however, parliament in April 1997 voted overwhelmingly to threaten Russia with loss of its military bases, should it fail to extend Russian military control over the separatist region. In 1998, the U.S. and Britain began an operation to remove nuclear material from Georgia, dangerous remains from its Soviet years. A darling of the West since his days as the Soviet Union's foreign minister, Shevardnadze was viewed far less favorably by his own people, who were frustrated by unemployment, poverty, cronyism, and rampant corruption. In the 2000 presidential elections, Shevardnadze was reelected with 80% of the vote, though international observers determined the election was marred by irregularities.
In 2002, U.S. troops trained Georgia's military in antiterrorism measures in the hopes that Georgian troops would subdue Muslim rebels fighting in the country. Tensions between Georgia and Russia were strained over the Pankisi Gorge, a lawless region of Georgia that Russia said had become a haven for Islamic militants and Chechen rebels.
In May 2003, work began on the Georgian section of the enormously ambitious Baku-Tbilisi-Ceyhan oil pipeline, which runs from Azerbaijan through Georgia to Turkey. The pipeline opened in July 2006.
Massive demonstrations began after the preliminary results of the Nov. 2003 parliamentary elections. The opposition party (and international monitors) claimed that the elections were rigged in favor of Shevardnadze and the political parties who supported him. After more than three weeks of massive protests, Shevardnadze resigned on Nov. 30. Georgians compared the turn of events to Czechoslovakia’s “velvet revolution.” In Jan. 2004 presidential elections, Mikhail Saakashvili, the key opposition leader, won in a landslide. The 36-year-old lawyer built his reputation as a reformer committed to ending corruption, and in his first two years as president, Saakashvili made significant progress in rooting out the country's endemic corruption. Saakashvili's ongoing difficulty has been reining in Georgia's two breakaway regions, South Ossetia and Abkhazia, both of which are strongly supported by neighboring Russia.
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